Disturbing, the mechanismis are really smart, with the path that could be activate at certain point, but i can't understand the link with beatle (but i'm french maybe did i miss something ;) )
Maybe having a more graph theme (computer, map).
The main issu is about understanding how it works, and why i die sometimes.
Actually there is an opposition, the hand made graphics create something poetic, and smooth, but the gameplay is really fast, time based, arcade.
Actually i think that with an other background, or maybe an other graphics, the game could have a more readable path for each level, and also be more consistent.
Actually i think that your game could be really better with an orthographic view, like Monument Valley. It could explain the lack of lisibility in your game path and could maybe became more poetic (if graphic like Monument Valley), or arcade oriented(like any retro aracde game but with beetle) :D
But it's still pretty fun, and easy to play.